[1] Western Gen Hosp, Chromosomes & Gene Express Sect, MRC Human Genet Unit, Edinburgh EH4 2XU, Midlothian, Scotland
[2] Univ Penn, Sch Med, Dept Cell & Dev Biol, Philadelphia, PA 19104 USA
[3] Univ Penn, Sch Med, Inst Regenerat Med, Epigenet Program, Philadelphia, PA 19104 USA